Marine biology, oyster biology, oyster aquaculture and hatcheries invertebrate pathology, environmental impact assessment and coastal zone management.
A.A., Mississippi Delta Junior College, 1940
Pharmacist's Mate I/C, U.S. Navy, 1942-1945
B.S. (Zoology) - Louisiana State University, 1942
M.A. (Biology) - Rice University, 1952
Ph.D., (Biology), Rice University, 1954